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Classification and evolution

  • Taxonomic groups = hierarchial groups in classification
  • Kingdom, Phylum, Class. Order, Family, Genus, Species
  • Recently, scientists have added another kingdom and 3 domains
  • Why use it?
    • identify species
    • predict characteristics
    • find evolutionary links
  • Binomical nomenclature --> Genus species (Canis familiaris)
  • Five kingdoms
    • Prokaryotae --> bacteria
      • unicellular 
      • no nucleus or membrane-bound organelles
      • circular DNA
      • no visible feeding mechanism
    • Protoctista --> amoeba
      • mostly unicellular
      • nucleus and membrane-bound organelles are present
      • some have chloroplasts
      • some have cilia or flagella
    • Fungi --> mushrooms, moulds or yeast
      • unicellular or multicellular
      • nucleus and membrane-bound organelles are present
      • cell wall = chitin
      • no chloroplasts
      • nutrients acquired from dead or decaying material
    • Plantae 
      • multicellular
      • nucleus, membrane-bound organelles and chloroplasts present
      • gametes move with cilia or flagella
      • autotrophic feeders
      • cell wall = cellulose
      • food stored as starch
    • Animalia 
      • multicellular
      • nucleus and membrane-bound organelles present
      • no chloroplasts or cell walls
      • hetereotrophic feeders
      • food stored as glycogen
  • Evolutionary relationships
    • as they become different their DNA changes so their proteins and sequence of amino acids also change
  • The 3 domains
  • Based on differences in the sequence of nucleotides in rRNA and the cell membrane lipid structures  and sensitivity to antibiotics
    • Eukarya
      • 80S ribosomes
      • RNA polymerase has 12 proteins
    • Archaea
      • 70S ribosomes
      • RNA polymerase has 8-10 proteins
    • Bacteria
      • 70S ribosomes
      • RNA polymerase has 5 proteins
  • Under this system, the Prokaryotae kindgom is split into Archaebacteria and Eubacteria
    • Archaebacteria
      • "ancient" bacteria
      • live in extremes like thermal vents
    • Eubacteria
      • "true" bacteria
      • found in all environments
  • Phylogeny = evolutionary relationships between organisms
  • Phylogenetic trees show the relationships with branches from the common ancestor
    • relies on physical characteristics and genetic makeup
    • can be done without reference to classification
    • continuous tree --> no groups means there's no species that there's debate over where they belong 
  • Evidence for evolution
    • Palaentology --> study of fossils
      • bacteria in the oldest rocks and invertebrates and vertebrates in the more recent ones which supports evolution because there's time between the lesser and more complex organisms
      • sequence in which organisms are found matches with ecological links (plants older than animals)
      • similarities in anatomy of fossils shows how closely related organisms  evolved from common ancestor
      • BUT fossil record isn't complete
    • Comparative anatomy --> similarities and differences in the anatomy of organisms
      • used because the fossil record isn't complete
      • homologous structure = structure that appears superficially different in different organisms or has a different function
        • provides evidence for divergent evolution (species have evolved from a common ancestor into different forms)
    • Comparative biochemistry --> similarities and differences in the proteins and other essential molecules
      • some molecules are highly conserved so changes can show evolutionary links
      • mostly study cytochrome c (protein in respiration) or rRNA
        • neutral evolution = most of the variability of the molecule occurs outside of its functional region so accumulation isn't affected by natural selection therefore changes to these regions can show how close two species are
        • the number of differences between a molecule is plotted against the rate at which the molecule undergoes neutral base pair substitutions so a point can be estimated where they last shared a common ancestor
        • rRNA is used because it has a slow rate of substitution 
  • Variation
  • interspecific variation = variation between members of different species
  • intraspecific variation = variation between members of the same species
  • causes = genetic or environmental
    • Genetic variation
      • alleles 
      • mutations
      • meiosis --> independent assortment and crossing over
      • sexual reproduction
      • chance
    • Environmental variation
      • plants are more affected than animals because they cannot move
      • sunlight
      • water availability
      • scars on the skin
    • Both
      • tall parents --> you have the alleles to be tall but if your diet isn't right or you suffer from a disease, it means you may grow below average height
      • skin colour --> determined by melanin in your skin, birth skin colour is purely genetic but the more you're exposed to sunlight, the concentrations of melanin change 
  • Discontinuous variation = when variation can only result in certain values --> blood type and bacterial cell shape (spherical, rods, spiral, comma or corkscrew)
    • controlled by a single gene
    • represented in a pie chart or bar chart
  • Continuous variation = when variation can be any value within a range --> height, mass
    • controlled by more than one gene as well as environmental factors
    • represented in line graphs or histograms
  • Normal distribution = bell-shaped curve
    • mean, mode and median are all the same
    • distribution is bell shaped or symmetrical around the mean
    • 50% are less than the mean and 50% are more than the mean
    • most values lie close to the mean
  • Standard deviation = how spread out the data is
  • Greater the SD, the greater the spread
  • Characteristic with a high SD has a large amount of variation
    • normal spread = 68% within 1 SD, 95% within 2 SD, 99.7% within 3 SD
  • Student's t test = compare the mean values of two sets of data
    • the data must be normally distributed and there muse be enough data to calculate a reliable mean
  • Spearman's rank correlation coefficient = used to consider the relationship between two sets of data
    • no correlation = no relationship
    • negative correlation = as one set increases, the other decreases
    • positive correlation = as  one set increases, so does the other
  • Adaptations
  • Anatomical adaptations = physical features
    • body covering = hair, scales, spines, feathers, shells to stay warm or protect them
    • camouflage = outer colour blends into environment to make it harder to predators or prey to spot you
    • teeth = shape and type are related to diet
    • mimicry = copying another animal's appearance or sounds to allow a harmless animal to fool predators
    • Marram grass --> curled leaves to minimise the surface area, hairs on inside surface area of the leaves to trap moist air, sunken stomata, thick waxy cuticle
  • Behavioural adaptations = the way the organism acts
    • survival behaviours = opossums play dead
    • courtship = elaborate courtship rituals to attract a mate
    • seasonal behaviour = migration to nesting sites or hibernation through the winter
    • innate = inherited through genes, doesn't have to be learned
    • learned = adaptations learned from observing other animals
  • Physiological adaptations = processes that take place inside the animal
    • poison production 
    • antibiotic production
    • water holding 
  • Anatomical adaptations provide evidence for convergent evolution
    • analogous structure = adapted to perform the same function but have a different genetic origin
    • convergent evolution takes place when unrelated species begin to share similar traits these are due to them evolving in similar environments
      • marsupial and placental mice - both are small and agile climbers that forage at night for small food items
      • flying phalangers and flying squirrels - both are gliders that eat insects and plants
      • marsupial and plancental moles - both burrow to find worms and grubs, they have streamlined bodies
  • Natural selection
    1. organisms within a species show genetic variation
    2. organisms who are best adapted to a selection pressure have an increased chance of survival and successfully reproducing --> survival of the fittest
    3. successful organisms pass on the allele encoding for the advantageous characteristic
    4. the process is repeated for each generation so over time the proportion of individuals with the advantageous allele increases in the gene pool
    5. over a long time the process can lead to speciation
  • Modern examples of evolution 
    • antibiotic-resistant bacteria --> MRSA
    • peppered moths --> industrial revolution meant that the black ones became camouflaged where the white ones didn't
    • sheep blowflies became resistant to the pesticide used against them
    • falvobacterium has evolved to digest nylon which is beneficial to humans as it clears up factory waste (developed nylonases due to gene replication combined with frameshift)

Biodiversity

  • Importance = essential for maintaining a balanced ecosystem for all organisms --> they provide food, oxygen and other materials for us to survive
  • Measuring a baseline for the biodiversity of an area means you can work out how much of an effect a change has had/ will have on the area
  • Habitat biodiversity = number of different habitats found within an area
    • each habitat will have different numbers of different species so the higher the habitat diversity the higher the biodiversity
  • Species biodiversity has two compononents:
    • ​​​​​​​species richness --> number of different species living in a particular area
    • species evenness --> comparison of the number of individuals of each species in a community
    • overall species diversity can be different even if it has the same number of species, it also depends on the evenness
  • Genetic biodiversity = variety of genes that make up a species
    • many of the genes are the same for all individuals in the species but alleles do exist and they cause the genetic variation within and between species
    • genetic biodiversity can result in different characteristics being exhibited (dogs can be big or small, round or pointy eared) 
    • greater genetic biodiversity within a species allows for better adaptation to a changing environment
  • Sampling = taking measurements of a limited number of individual organisms within a particular area
  • Used to estimate the number of organisms in an area aka the abundance of the organisms
  • Also used to measure a particular characteristic of an organism (average height of crop)
  • Random sampling = selecting individuals by chance
    • each population has equal chance of being recorded
    1. mark out a grid on the grass using tape measures
    2. use random numbers to determine coordinates
    3. take a sample from each randomly generated coordinte pair
  • Non-random sampling = sample is not chosen at random
    • opportunistic --> weakest form of sampling as it is not representative of the whole population (uses organisms that are conveniently available)
    • stratified --> some population can be divided into sub-groups based on a particular characteristic, a random sample is then taken from each of the groups proportional to its size
    • systematic --> areas are identified within a habitat that are then sampled separately --> usually with line (line along the ground and take samples from specified points) or belt transect (two parallel lines marked and samples taken in the area between them)
  • Reliability
    • Sample is never entirely representative because:
      • sampling bias --> may be by accident or deliberate (you choose an area because you know it'll give you good numbers for certain species)
      • chance --> the organisms selected may not be representative of the whole population --> can never be removed but its effects can be minimised with large sample sizes 
  • Sampling animals
    • pooter to catch insects
    • sweep nets to catch insects in long grass
    • pitfall traps for small invertebrates
    • tree beating for invertebrates that live in trees or bushes (shake it over a white cloth to catch them)
    • kick sampling for organisms in a river (the riverbed is "kicked" for a while before to disturb the substrate then a net catches anything that's disturbed downstream)
  • Estimating animal population size
    • capture-mark-release-recapture technique
    • capture as many as you can
    • mark them all and then release them
    • time is allowed for the organisms to redistribute themselves
    • repeat the capture process and compare the numbers that are new or recaptured
    • the greater the marked individuals recaptured the smaller the population size
  • Sampling plants
    • point quadrat --> frame with a horizontal bar, at set intervals a pin is pushed through to the soil and each plant that touches the pin is recorded
    • frame quadrat --> sqaure frame divided into a grid, the type and number of species and individuals within each section is recorded
      • used to measure:
      • density --> if plants can be seen clearly, counting how many are in one quadrat shows the density of the plant per metre <-- not an estimate
      • frequency --> used where individuals of a species are hard to count --> count the number of squares the species is present within to work out an estimated percentage (clover in 65 of 100 squares means they are in 65% of the area)
      • percentage cover --> used for speed when collecting data --> used when a species is abundant but hard to count --> estimate by eye of the area within the quadrat that the plant is present within
    • for best results, quadrats should be used in random sampling but often aren't)
  • Measuring species richness
    • make a list of all the species identified then a total number can be calculated
    • identification keys are used when sampling
  • Measuring species evenness
    • compare the numbers for the individuals in each species
  • Measuring abiotic factors
    • abiotic factors = non living components in a habitat
    • wind speed
    • light intensity --> light meter
    • relative humidity --> humidity sensor
    • pH --> pH probe
    • temperature --> temperature probe
    • oxygen content of water --> dissolved oxygen probe
    • can be measured quickly and accurately so that:
      • rapid changes can be detected
      • human error in taking reading is reduced
      • high degree of precision
      • data can be stored and tracked
  • Calculating biodiversity
    • use the Simpson's Index of Diversity to take both species evenness and richness into account
    • values always between 1 and 0 --> closer to 1 means higher biodiversity
    • organisms in a low biodiversity habitat are usually highly adapted to survive there
  • Genetic diodiversity
    • differences in alleles create genetic biodiversity
    • species with greater genetic biodiversity are more likely to adapt and survive as it is likely that some organisms carry the advantageous alleles
    • increased by:
      • mutations in the DNA
      • interbreeding between different populations as the alleles are transferred between the groups aka gene flow
    • decreased by:
      • selective breeding where organisms are only selected to breed if they have desirable traits
      • captive breeding programmes in zoos and conservation centres as new alleles are rarely introduced
      • rare breed where they have been selectively bred but then go out of fashion so numbers and allele numbers drop dramatically
      • artificial cloning
      • natural selection --> organisms evolve to only carry the advantageous alleles
      • genetic bottlenecks --> few individuals survive an event or change so the gene pool is reduced
      • genetic drift due to random nature of alleles being passed from parent to offspring (more pronounced in populations with small gene pools)
  • Measuring genetic biodiversity
    • measure polymorphism (polymorphic genes have more than one allele like immunoglobulin which codes for blood type)
    • most genes are monomorphic where only a single allele exists (ensure the basic stucture of a species stays the same)
    • proportion of genes that polymorphic can be measured by:
      • number of polymorphic gene loci / total number of loci
    • the greater the proportion of polymorphic genes, the greater the genetic biodiversity in a population
  • Factors affecting biodiversity
    • deforestation
      • can occur naturally but it is mostly deliberate by humans
      • affects biodiversity by:
      • directly reduces the number of trees in an area
      • specific species are sometimes targeted so it reduces species biodiversity
      • destroys the habitats for animals so their numbers fall too
      • animals are forced to migrate to ensure their survival
    • agriculture 
      • selection of only a few species reduces the biodiversity of the area
      • much of the methods used by farmers reduces biodiversity:
      • deforestation to create area for planting 
      • removal of hedgerows
      • use of pesticides and herbicides
      • monoculture 
    • climate change and global warming
      • melting of the polar ice caps would lead to the extinction of the plants and animals in these regions
      • rising sea levels from the melting caps and thermal expansion of the seas would lead to flooding
      • higher temperatures and less rainfall would result in droughts and some plant species dying out + xerophytes would become more dominant --> less food for some animals so the ones who eat xerophytes would be more common
      • insect life cycles and populations would change as they adapt to the new climates --> as key pollinators they would effect the plants as well
      • slow climate change would give time for species to adapt and as this would lead to the loss of native species, biodiversity wouldn't be lost
  • Reasons for maintaining biodiversity
    • aesthetic reasons:
      • different plants and animals enrich out environments and lives
      • natural world provides inspiration for the arts
      • patients recover more quickly from stress if they're surrounded by plants
    • economic reasons:
      • soil erosion may occur as a consequence of deforestation so there would be a drop in land available for crops and animals
      • important to conserve the organisms we use to make things (hardwood timber) to prevent industries breaking down
      • protect species with potential economic importance --> undiscovered medicines
      • continuous monoculture depletes the soil which would lead to smaller crop yields and less money
      • high biodiversity protects against abiotic stresses so crops are more likely to survive
      • economic income from areas of natural beauty
      • plant varieties are needed for cross-breeding to make more profitable and successful crops
    • ecological reasons:
      • the removal of one species could have a serious effect on the others in its ecosystem - including us!
      • keystone species have a disproportionally large effect on their environments so its essential to protect them so other species don't also disappear
  • Human activity vs biodiversity
    • in many countries the natural habitat is created and maintained by human intervention by farmers and landowners
  • Maintaining biodiversity
  • In situ conservation = within the natural environment
    • maintains the genetic diversity and the evolutionary adaptations
    • preserves interdependent relationships therefore interlinked species are also protected
    • wildlife reserves:
      • controlled grazing
      • restricted human access
      • controlling poaching
      • feeding animals
      • reintroduction of a species
      • removal of invasive species
      • halting succession
    • marine conservation zones:
      • less well established
      • vital in preserving species-rich areas
      • aim is to create areas of refuge where populations can build up and repopulate adjacent areas
  • Ex situ conservation = outside of the natural environment
    • botanic gardens:
      • plant species actively maintained to provide optimum conditions
      • roughly 1500 worldwide holding 35,000 plant species
      • majority of species aren't conserved though
      • many wild species are unrepresented
    • seed banks:
      • store of genetic material
      • seeds are carefully stored so new plants can be grown in the future
      • dried and stored at -20C
      • don't work for all plants, some seeds die when dried and frozen (mostly the seeds from rainforests)
    • captive breeding programmes:
      • often run and managed by zoos and aquatic centres
      • scientists help to create a stable, healthy population of the species so it can be gradually reintroduced to its natural habitat
      • provide the animals with shelter, food, no predators and vets
      • suitable breeding partners or semen can be imported from other zoos to increase the gene pool
      • maintaining genetic biodiversity can be difficult as interbreeding happens often
      • sometimes they cant't be reintroduced because:
        • loss or resistance to a disease
        • may not have learned the correct behaviours to survive
        • genetic make up of the captive species may be too different from wild ones
        • in many cases the natural habitat must first be restored
    • conservations agreements
      • International Union for Conservation of Nature (IUCN) assist in securing agreements between nations
      • established the Convention of International Trade in Endangered Species (CITES) which regulates the trade of wild plant and animal specimens (protects 35,000 species)
      • the Rio Convention of 1992 --> 172 nations agreed on :
        • Convention of Biological Diversity (CBD) requires them to develop national strategies for sustainable development
        • United Nations Framework Convention on Climate Change (UNFCCC) agreed to take steps to stabilise greeenhouse gases
        • United Nations Convention to Combat Desertification (UNCCD) aims to prevent desertification of fertile land

Classification and evolution

Sara Bean
Module by Sara Bean, updated more than 1 year ago
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